7-Day Western Islands: Fernandina's Marine Iguanas

Most Galapagos itineraries begin here. Baltra's airport is the world's first fully ecological airport, run on solar and wind power and built largely from recycled materials, including old oil pipes repurposed into the terminal structure. Land iguanas, wiped out here by the 1950s and reintroduced through a decades-long breeding program, now number in the hundreds again, some right around the airport grounds. From the terminal, a short bus transfer brings you to the pier for the panga ride out to your ship.

Up in the cool, misty highlands of Santa Cruz, a short walk brings you among wild giant tortoises moving through the undergrowth on their own terms. No fences, no enclosures, just several-hundred-pound reptiles going about their day. This is one of the best places anywhere in the islands for that encounter, and the surrounding vegetation is good habitat for several of Darwin's finches, heard as often as seen.

Sierra Negra's caldera stretches 10 km north to south and 9 km east to west, one of the largest volcanic calderas on Earth. The more commonly walked route to Volcan Chico crosses relatively recent lava flows with sweeping views toward the volcanoes of Isabela and neighboring Fernandina; a longer trek reaches a system of active fumaroles known as the Sulfur Mine. This is a geology-and-scenery visit, and wildlife isn't the focus here.

At first glance Punta Moreno looks like little more than a lava flow at the base of Cerro Azul, but its black surface holds a series of lagoons with a surprising amount of life. Flamingos are a fairly good bet in the pools, and paint-billed crakes, white-cheeked pintails, and common gallinules turn up for those willing to watch the water closely.

A zodiac ride along Tagus Cove's cliffs is the first stop, tracing walls carved with the graffiti of sailors and whalers going back over a century. Charles Darwin himself anchored here in 1835. Watch the rock for Galapagos penguins and flightless cormorants before you land and set off on a gently climbing trail, roughly 30 minutes each way, to a viewpoint over Darwin Lake, a crater lake saltier than the ocean itself. Along the way, look for land birds that are easy to miss: ground and tree finches, yellow warblers, large-billed flycatchers, and, with a little luck, the tool-using woodpecker finch. Land iguanas and giant tortoises sometimes cross the path as well. Back at the water, most visits close with a dinghy ride or a snorkel along the submerged cliff wall, depending on the day's plan, with another good chance at penguins and cormorants gliding by.

A wooden staircase of 372 steps climbs Bartolome's extinct volcano from a lava-and-concrete dock or sandy beach, depending on conditions. Along the way, red, orange, black, and even green volcanic layers mark the slopes, home to pioneer plants like the island-endemic Tiquilia nesiotica, lava cactus, and scattered Scalesia. With some luck, a lava lizard crosses the trail. At the top, the view over the twin bays and the iconic Pinnacle Rock is the reward for the climb. Back at sea level, a snorkel around the coral reef and submerged lava rocks is a good bet for Galapagos penguins and white-tipped reef sharks, with sea turtles and manta rays turning up now and then too.

One of the few visitor sites directly tied to an active conservation program, run jointly by the Charles Darwin Foundation and the Galapagos National Park, working to restore the local land iguana population. From the rocky landing, which doubles as a good snorkeling spot, a trail leads inland past two small saltwater lagoons, sometimes home to flamingos, before circling the hill itself, where land iguanas and their burrows dot the hillside.

A relaxed, sandy stop on Santa Cruz's north shore. A lagoon just behind the beach occasionally draws flamingos and other shorebirds, and the beach itself is one of the archipelago's main sea turtle nesting grounds, most active between December and June. It's an easy visit, as much about the swim as the wildlife.