8-Day Western Islands: Elizabeth Bay's Penguins

Most Galapagos itineraries begin here. Baltra's airport is the world's first fully ecological airport, run on solar and wind power and built largely from recycled materials, including old oil pipes repurposed into the terminal structure. Land iguanas, wiped out here by the 1950s and reintroduced through a decades-long breeding program, now number in the hundreds again, some right around the airport grounds. From the terminal, a short bus transfer brings you to the pier for the panga ride out to your ship.

A relaxed, sandy stop on Santa Cruz's north shore. A lagoon just behind the beach occasionally draws flamingos and other shorebirds, and the beach itself is one of the archipelago's main sea turtle nesting grounds, most active between December and June. It's an easy visit, as much about the swim as the wildlife.

You won't set foot on Daphne Major. Steep cliffs ring almost the entire island, and it's closed to tourists, reserved for scientists with a special research permit. What makes it worth pointing out as you cruise past is the science: this tiny volcanic cone, barely half a mile long, is where Peter and Rosemary Grant spent over 40 years documenting Darwin's finches in real time, watching natural selection reshape beak size from one generation to the next. It is the research behind the Pulitzer-winning book The Beak of the Finch.

A panga brings you ashore onto one of the most distinctive beaches in the islands, with sand tinted deep red by its high iron content, with Rabida's rugged cliffs rising behind it. Sea lions are usually stretched out along the shoreline, and it's a good spot to watch for the endemic Galapagos dove working the sand nearby. From the beach, a short trail winds through palo santo forest and cactus scrub, climbing gently to an overlook with sweeping views across the bay, a nice place to pause and take in the color of the landscape before heading back down. If you'd rather stay on the water, snorkeling right off the beach is an easy, relaxed way to spend the visit, with good odds of company from the resident sea lions.

There's no landing at Punta Vicente Roca: this dramatic, eroded caldera on Isabela's coast is explored entirely by snorkel, dinghy, or kayak. The cold Cromwell Current runs through here, and it's what keeps the water so alive: blue-footed boobies nest reliably along the cliffs, and in the water you can expect Galapagos penguins and, between July and November, ocean sunfish rising to cleaning stations in the cold upwelling. Sperm whales pass through on rare occasions, and giant manta rays are a possibility during the same cooler months.

At first glance Punta Moreno looks like little more than a lava flow at the base of Cerro Azul, but its black surface holds a series of lagoons with a surprising amount of life. Flamingos are a fairly good bet in the pools, and paint-billed crakes, white-cheeked pintails, and common gallinules turn up for those willing to watch the water closely.

At the base of Alcedo Volcano on Isabela, Urbina Bay marks a dramatic geological event: a 1954 uplift left blocks of coral fully exposed on dry land, well above the waterline. A long, level walk leads out to them, passing land iguanas and flightless cormorants along the way, and, on occasion, a giant tortoise.

A quiet landing on Santiago's western coast, best known as a nesting ground for Pacific green sea turtles, who come ashore to lay their eggs between December and June.

A dry landing sets you down on North Seymour's flat, sun-baked terrain, and the trail wastes no time getting to the action: magnificent frigatebirds display from low shrubs right along the path, males inflating their scarlet throat pouches into balloons to catch a female's eye. Blue-footed boobies are a near-constant presence along the same stretch, and during the breeding season you may catch their high-stepping courtship dance just a few feet from the trail, showing off the bright blue feet that give the species its name. Land and marine iguanas share the coastline as the trail loops back toward the water, and sea lions haul out along the shore. Keep an eye on the cliffs above the surf too, where swallow-tailed gulls sometimes perch between flights out to sea.

Set on the grounds of the Galapagos National Park Service on Santa Cruz, this station was once home to Lonesome George, the last known tortoise of the Pinta subspecies. A series of interpretive buildings walks you through the conservation science underway across the islands, and the surrounding native vegetation is a good place to look for Darwin's finches going about their business.