6-Day Southern Islands: Floreana's Devil's Crown

San Cristobal's airport sits at the edge of Puerto Baquerizo Moreno, a five-minute drive from the town center, so this is usually your first taste of the islands rather than a standalone excursion. Sea lions and marine iguanas are known to rest right next to the runway here, an unscripted welcome few airports can match. From the terminal, a short transfer takes you down to the pier for a panga ride out to your ship.

Named for a park ranger who dedicated 39 years of service to the islands, this San Cristobal facility focuses on conserving the island's native Chelonoidis chathamensis giant tortoise. Established in 2002, it lets you see tortoises at every stage of growth in a carefully designed, semi-natural habitat, part of an ongoing breeding and reintroduction program.

A dry landing among the rocks puts you right at the start of one of the richest walks in the islands. The trail follows dramatic cliffs where blowholes send spray high into the air with every swell, and along the way marine iguanas bask on the black rock. Between December and March, the males here turn a striking red and green for the breeding season. Punta Suarez is the only place on Earth where waved albatrosses nest, and between April and December you'll find the colony in full swing, their courtship displays peaking in October. Blue-footed and Nazca boobies nest along the same stretch of cliff, red-billed tropicbirds circle overhead, and the endemic Espanola mockingbird, famously bold, often comes right up to visitors. Keep an eye out too for the Galapagos hawk and the Galapagos dove working the scrub beside the trail.

A short landing on one of the most beautiful beaches in the islands brings you into a resident colony of Galapagos sea lions, sprawled across the white sand. Between August and November you may also spot pups nursing nearby. This is one of the only places to see the Espanola mockingbird, found nowhere else on Earth and famously unbothered by visitors. After time on the beach, a relaxed snorkel just offshore is a good way to close out the visit, with spotted eagle rays often gliding beneath the surface.

From the landing, a trail leads to a wide, shallow lagoon where Galapagos flamingos wade through the shallows, filtering shrimp from the mud that gives them their color. Continuing over a low rise brings you to Flour Beach, a nesting ground for green sea turtles between December and June. Just offshore, the eroded remains of an old volcanic cone form Devil's Crown, one of the best snorkeling spots in the islands. White-tipped reef sharks and spotted eagle rays are a near-certain sight inside the ring, and on a good day you might also spot hammerhead sharks, sea lions, or a colorful mix of reef fish weaving through the rocks.

From the landing, a trail leads to a wide, shallow lagoon where Galapagos flamingos wade through the shallows, filtering shrimp from the mud that gives them their color. Continuing over a low rise brings you to Flour Beach, a nesting ground for green sea turtles between December and June. Just offshore, the eroded remains of an old volcanic cone form Devil's Crown, one of the best snorkeling spots in the islands. White-tipped reef sharks and spotted eagle rays are a near-certain sight inside the ring, and on a good day you might also spot hammerhead sharks, sea lions, or a colorful mix of reef fish weaving through the rocks.

A scenic lookout point on Floreana overlooking the water below, where brown pelicans, red-footed boobies, magnificent frigatebirds, and the diminutive Elliot's storm-petrel are often seen over the surrounding cliffs and sea.

One of the few places in the islands where human history takes center stage over wildlife. In 1793, a whaling captain installed a wooden barrel here as an informal mailbox, and the tradition has never stopped: you're welcome to leave a postcard without a stamp, and to sort through what's already inside for anything addressed near your own home to hand-deliver in person. After a wet landing on the sandy beach, a short walk brings you to the barrel. It is one of the most memorable traditions anywhere in the islands.

Formed by geological uplift rather than volcanic activity, Santa Fe is unusually flat for a Galapagos island, and it's the only place on Earth with its own endemic subspecies of land iguana. After landing in clear, turquoise water, a walk through a forest of giant, tree-sized Opuntia cactus leads to iguanas basking beneath them. The Galapagos dove is a reliable sight along the trail, and with some luck you may also spot the Galapagos snake or the Galapagos hawk.

A short landing puts you immediately among sea lions and the endemic Plaza land iguana, smaller and more golden than its relatives elsewhere. Watch closely and you might spot a rare hybrid, the result of an occasional cross between a marine and a land iguana found only here. The trail crosses a carpet of succulent Sesuvium, red for part of the year, studded with Opuntia cactus. Along the cliffs, swallow-tailed gulls nest on the ledges beside red-billed tropicbirds, and frigatebirds circle overhead as the loop returns to the landing.

A dry landing sets you down on North Seymour's flat, sun-baked terrain, and the trail wastes no time getting to the action: magnificent frigatebirds display from low shrubs right along the path, males inflating their scarlet throat pouches into balloons to catch a female's eye. Blue-footed boobies are a near-constant presence along the same stretch, and during the breeding season you may catch their high-stepping courtship dance just a few feet from the trail, showing off the bright blue feet that give the species its name. Land and marine iguanas share the coastline as the trail loops back toward the water, and sea lions haul out along the shore. Keep an eye on the cliffs above the surf too, where swallow-tailed gulls sometimes perch between flights out to sea.

A relaxed, sandy stop on Santa Cruz's north shore. A lagoon just behind the beach occasionally draws flamingos and other shorebirds, and the beach itself is one of the archipelago's main sea turtle nesting grounds, most active between December and June. It's an easy visit, as much about the swim as the wildlife.

A tiny, flat islet of white coral sand between North and South Seymour, Mosquera is best known for its resident sea lion colony, sprawled across the beach for much of the day. Shorebirds and lava herons work the shoreline fairly reliably, and sharks and rays are a possibility for anyone snorkeling or diving nearby.

The return transfer is straightforward: a panga ride from your ship to the pier, then a short bus ride to Baltra's airport, the world's first fully ecological terminal, run on solar and wind power. Land iguanas, reintroduced here after being wiped out decades ago, are sometimes seen right around the grounds on the way to your flight.